①CAS Member, Institute of Neuroscience, State Key Laboratory of Neuroscience, Shanghai Institutes for Biological Sciences, CAS,Shanghai 200031, China; State Key Laboratory of Brain and Cognitive Science, Institute of Biophysics, CAS, Beijing 100101, China; ②③④Ph.D., Institute of Neuroscience, State Key Laboratory of Neuroscience, Shanghai Institutes for Biological Sciences, CAS,Shanghai 200031, China