自然杂志 ›› 2026, Vol. 48 ›› Issue (1): 9-17.doi: 10.3969/j.issn.0253-9608.2026.01.002

• 特约专稿 • 上一篇    下一篇

中国化石见证的马类演化史

邓涛   

  1. 中国科学院古脊椎动物与古人类研究所,北京 100044
  • 收稿日期:2026-01-15 出版日期:2026-02-25 发布日期:2026-02-06
  • 基金资助:
    国家自然科学基金重点项目(42430207)

The evolutionary history of horses witnessed by Chinese fossils

DENG Tao   

  1. Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing 100044, China
  • Received:2026-01-15 Online:2026-02-25 Published:2026-02-06

摘要: 中国广袤的土地上保存了跨越5 600万年的马类化石序列,构建了从起源、辐射到现生类群崛起的完整全球演化链条,提供了无可替代的东方证据。湖南衡阳早始新世的丁氏曙马填补了亚洲早期马类化石记录的空白,这一发现将马类在亚洲的历史追溯到起源阶段,并揭示了其可能经过欧洲-北美-亚洲的早期洲际迁徙路径,修正了认为马类仅源于北美和欧洲的传统观点。文章重点阐述了中新世中国马类演化的独特图景,集中体现在中华马与三趾马的共生演化,二者在中国北方同一地层中共存,反映了森林-草原混合景观。通过碳氧同位素等方法,证实以长鼻三趾马为代表的三趾马才是最早、最适应开阔草原的马类,而非其后的真马。此外,青藏高原的三趾马化石成为重建古高度的关键证据。甘肃临夏盆地发现的埃氏马是东亚最早的真马,与三趾马的共生记录了演化的世代交替。随后,中国特有的三门马、庆阳马等展现了真马的快速辐射。尤其关键的是,周口店三门马和北京马作为北京猿人的伴生动物,其大量化石见证了早期人类对马类的狩猎活动,即人类活动开始介入马类演化的历史。

关键词: 马类, 演化, 地层, 气候环境, 新生代, 中国

Abstract: China’s vast land preserves a sequence of horse fossils spanning 56 million years, constructing a complete global evolutionary chain from their origin, radiation to the rise of extant taxa, providing irreplaceable oriental evidence. The Early Eocene Erihippus tingae from Hengyang, Hunan, fills the gap in the early equid fossil record of Asia. This discovery dates the history of equids in Asia back to the origin stage and reveals a possible early intercontinental migration path from Europe via North America to Asia, revising the traditional view that equids originated only in North America and Europe. The unique picture of equid evolution in China during the Miocene is highlighted, particularly reflected in the symbiotic evolution of Sinohippus and Hipparion, which coexisted in the same strata in northern China, reflecting a mixed forest-steppe landscape. Through methods such as carbon and oxygen isotopes, it is confirmed that the three-toed horse represented by Proboscidipparion is the earliest and most adapted to open grasslands, rather than the later true horses. In addition, the Hipparion fossils from the Tibetan Plateau became key evidence for reconstructing paleoelevation. Equus eisenmannae discovered in the Linxia Basin, Gansu, is the earliest true horse in East Asia, and its symbiosis with the three-toed horse records the generational alternation of evolution. Subsequently, the unique Chinese horses such as E. sanmeniensis and E. qingyangensis exhibited rapid radiation of true horses. Especially crucial are E. sanmeniensis and E. beijingensis from Zhoukoudian as companion animals of Peking Man, which have a large number of fossils witnessing early human hunting activities on horses, indicating the beginning of human intervention in the history of equid evolution.